How can I find out which command/action is executed on a shortcut combination?

I have created a custom shortcut and bound the shutter -s to it:

Then I tried to find out where this command is stored and checked settings by dconf-editor. But there are no information about the shutter -s bound command and Shift+Print combination:

Thus, the questions:

1. Where full information about key bindings is stored? Does a way exist to say: this key combination is using this command? For example, I want to see the default screenshot command which is executed on the Print press.
2. What happens when I press Print key? How this key press achieves a screenshot utility? For example:
   • some tty gets this key
   • GNOME Shell is connected to this tty, so it gets the key as input.
   • then GNOME Shell looks at some database (dconf?) and executes the command bound to the Print key.

I have read similar questions, like

• How can I find which command is bound to a given keyboard shortcut?
• Where are GNOME keyboard shortcuts stored?

but haven't found an answer to my questions.